www.CuriousTaxonomy.net
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Curiosities of Biological Nomenclature
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Mark Isaak
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Etymology: Names from Fictional Characters
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Antigone Reichenbach, 1853 (crane)
Crito Distand, 1916 (leafhopper) named after
a dialog by Plato.
Electra Lamouroux 1816 (bryozoan)
Gargantua Jullien, 1888 (bryozoan)
Gargantuavis philoinis Buffetaut and Le Loeuff, 1998
(huge Cretaceous flightless bird) This French fossil was named for one
of the giants in Rabelais'
Gargantua and Pantagruel. "Philoinis" means "wine-loving", which
describes the original Gargantua but probably not the bird.
Grendelius McGowan, 1976 (Jurassic ichthyosaur) named
for Beowulf's nemesis. Now synonymized with
Brachypterygius.
Agra othello Erwin, 2000 (carabid)
Iago Compagno & Springer, 1971 (shark)
Oberonia Lindl. (1830) (fairy orchid) The
array of tiny flowers on a stalk connected to a cluster of leaves
evoked an image of a tiny carriage with a team of horses, suggesting
Oberon, king of the fairies, driving through the branches. The genus
Titania Endl. 1833 is now included
in Oberonia, and
Oberonia titania (Endl.) Lindl. pairs the
king and queen of the fairies.
Ophelia (annelid)
Peneothello (robin) probably so named
because the bird is mostly black ("pene" means "almost").
Pigrogromitus Calman, 1927 (sea spider),
and
Queubus Barnard, 1946 (sea spider) Both
species are named for imaginary characters fabricated by the Clown
character in Shakespeare's Twelfth Night (II, iii). In a
review of sea spiders ("The Pycnogonida of the Western North
Atlantic and the Caribbean," Proc. U.S. Nat. Mus. 97:
157-342, 1948), Joel Hedgpeth wrote, "In sooth, we taxonomists are
hard put to it to find names, but there have been far worse sources
than the nonsense of Will Shakespeare."
Puck (anglerfish)
Oedipus rex (salamander)
Oedipodrilus oedipus Holt (worm)
Saguinus oedipus oedipus (cotton-top
tamarin) According to a
paper
presented by A. J. Ginther and C. T. Snowdon at the 2004 American
Society of Primatologists conference ("The Oedipal conflict in
Saguinus oedipus"), these tamarins really do love their mothers
(though the dams do not let them complete the process). Apparently,
though, this behavior was not observed until after the species was
named, perhaps for its big feet.
Ozymandias Jordan & Gilbert, 1919 (fossil
fish)
Wukongopterus Wang et al., 2009 (Jurassic
pterosaur) Named for Sun Wukong, the Monkey King, hero of the Chinese
classic novel Journey to the West.
Agra eponine Erwin, 2000 (carabid) Named after the
street urchin in Les Miserables "who, in the Broadway version
of the story, personified tragic beauty. Such is the state of the
tropical forests where these beetles live."
Akela Peckham and Peckham, 1896 (jumping
spider) Named after a character in Kipling's The Jungle
Book.
Bagheera kiplingi Peckham and Peckham, 1896
(Central American spider) Named for Rudyard Kipling and Bagheera, the
black panther from Kipling's The Jungle Book. Ironically, it
is the first known vegetarian spider.
Anchylorhynchus pinocchio De Madeiros &
Nunez-Avellaneda, 2013 (weevil) because of its extremely
long snout as compared with others of its genus. (The genus name
translates as "crooked-snout".)
Litoria pinocchio Oliver et al. 2019
(treefrog) Males have "distintive fleshy rostral spikes."
[ZooTaxa 4604(2)]
Walckenaeria pinocchio Kaston, 1945 (spider) Named for
Carlo Lorenzini's prevaricating marionette in The Adventures of
Pinocchio.
Balnibarbi Fortey (trilobite) Named for the
abysmally inept technocracy in Swift's Gulliver's
Travels.
Viola lilliputana Ballard & Iltis, 2012
(violet) One of the smallest terrestrial dicots; named for the tiny
Lilliputians in Swift's Gulliver's Travels. [ZooKeys
430: 1]
Dryadella lilliputiana Cogniaux
(orchid)
Salticus lilliputanus (jumping
spider)
Peruphorticus gulliveri Erwin &
Zamorano (ground beetle) The beetle's large size relative to
others of its genus reminded the authors of Gulliver's travels in
Lilliput.
Borogovia Osmólska, 1987 (theropod
dinosaur) from "borogove", a mimsy creature from Lewis Carroll's poem
"Jabberwocky".
Daggoo, Queequeg, and
Tashtego Sime & Wahl, 2002 (ichneumonid
wasps) named for the harpoonists in Herman Melville's Moby Dick.
[Zool. J. Linn. Soc. 134: 1]
Albicetus Boersma and Pyenson, 2015
(Miocene whale) The genus name means "white whale", in reference to
Melville's Moby Dick. The authors do not know the whale's skin color,
but the bone is white, and the fossil, like its namesake, has a
deformed lower jaw. [PLoS ONE 10(12): e0135551]
Desulforudis audaxviator Chivian et
al. 2008 (sulfate-reducing bacterium) Found in water samples
from 2.8 km underground in the Mponeng gold mine in South Africa; the
only species known in its ecosystem. The name comes from a quotation
from Jules Verne's Journey to the Center of the Earth. The
hero, Professor Lidenbrock, finds a secret Latin inscription that
reads: "Descende, audax viator, et terrestre centrum attinges"
(Descend, bold traveller, and you will attain the center of the
Earth).
Dracula Luer 1978 (orchid) The orchid is blackish-red
and looks like a bat. The genus includes many species, including
Dracula vampira,
Dracula nosferatu, and
Dracula diabola. The species
D. fafnir is named for a Norse giant who
turned into a dragon to guard a horde of treasure.
(The genus name has also been used erroneously for the
pigeon genus Ducula.)
Liparis draculoides (another orchid)
Bradycneme draculae Harrison & Walker,
1975) (Cretaceous theropod)
Danionella dracula Britz (fish) a Burmese
fish with "fangs" made of bone.
Deinocroton draculi Penalver, et.al., 2017
(Cretaceous tick) Found in amber; its morphology suggests it lived in
nests of feathered dinosaurs. The genus means "terrible
tick."
Desmodus draculae Morgan, Linares and Ray, 1988 (giant
South American vampire bat, recently extinct).
Draculo Snyder, 1911 (dragonet fish)
Supraserphites draculi Rasnitsyn &
Öhm-Kühnle 2019 (Cretaceous wasp) Because of its large
mouthparts.
Hemignathus vorpalis James & Olsen 2003
(greater nukupu'u, an extinct Hawaiian finch) Named for the vorpal
blade in Lewis Carroll's poem "Jabberwocky", due to its long upper
bill.
Holorusia brobdingnagius (crane fly) Named for the
Brobdingnags, a race of giants in Swift's Gulliver's Travels.
According to Guinness, it is probably the world's largest crane fly, it
has a wingspan of 4 inches, and its legs may spread almost 9
inches.
Ichabodcraniosaurus Novacek 1996 [nomen
nudum] (dinosaur) Named for a character in Irving's Legend of Sleepy
Hollow. It was found without a head; a head was found later, but it
is uncertain whether the head belongs to the skeleton.
Agra ichabod Erwin, 2002 (carabid) "Refers
to the fact that the holotype is missing its head and the illusion is
that of the frightened schoolteacher Ichabod Crane's phantom nemesis,
the Headless Horseman, in 'The Legend of Sleepy
Hollow'...."
Laputa Whitley, 1930 (fish) and, more
appropriately,
Laputavis Dyke, 2001 (Middle Eocene fossil
swift) named for the floating castle in Swift's Gulliver's
Travels.
Messua Peckham and Peckham, 1896 (jumping
spider) Named after a character in Kipling's The Jungle
Book.
Millerocaulis tekelili Vera, 2012
(Cretaceous fern) "Tekeli-li" was a cry of the Tsalalians, an
Antarctic race in Edgar Allen Poe's The Narrative of Arthur Gordon
Pym of Nantucket. H. P. Lovecraft later make it an endlessly
repated cry of the monstrous shoggoths, which lived in Antarctic
caverns in At the Mountains of Madness. The fossil fern was
discovered in Antarctica. [Alcheringa 36: 37]
Morlockia Garcia-Valdecasas, 1984
(cave-dwelling remipede crustacean) Named for the Morlocks, the
subterranean subhumanoids in H.G. Well's The Time
Machine.
Muscatheres Evenhuis, 1986 (bee fly)
"There are only three Muscatheres known," referring to three specimens
of the lone species, M. lurida (previously described in the genus
Phthiria).
Nagaina Peckham and Peckham, 1896 (jumping
spider) Named after a character in Kipling's
"Rikki-Tikki-Tavi."
Paramphientomum yumyum Enderlein, 1907
(psocopteran) Probably named after the character Yum-yum in Gilbert
and Sullivan's "The Mikado." This is not confirmed, but it is
supported by the fact that the insect is native to Japan.
Phanuromyia princeps Nesheim and
Phanuromyia pauper Nesheim and Masner, 2017
(wasps) Names inspired by Mark Twain's The Prince and the
Pauper; P. pauper lacks longitudinal costae which
decorate P. princeps. [ZooKeys 663: 71]
Poanes hobomok (Harris, 1862) (Hobomok
skipper) Named for the Native American in Hobomok, a Tale of Early
Times an 1824 novel by Lydia Maria Child.
Pseudione quasimodo Boyko & Williams,
2004 (parasitic isopod (Crustacea: Isopoda: Bopyroidea) found
under the carapace of hermit crabs) Named for Victor Hugo's bell
ringer of Notre Dame, as the parasite has a distinct bulge in dorsal
view. (Boyko originally wrote in the manuscript that he "had a hunch"
it was a new species, but the un-amused editor insisted that be
stricken from the text.)
Semiramis Becker, 1913 (bombyliid fly) A story by
Voltaire about a Babylonian queen.
Sirenoscincus mobydick Miralles et al.,
2012 (skink) "The specific epithet refers to Moby Dick, the
famous albino sperm whale imagined by Herman Melville (1851), with
whom the new species shares several uncommon characteristics, such as
the lack of hindlimbs, the presence of flipper-like forelimbs, highly
reduced eyes, and the complete absence of pigmentation."
[Zoosystema 34: 701]
Stylaclista quasimodo Early (diapriid
wasp)
Tetragnatha quasimodo (Hawaiian spider)
Named for the kyphotic bellringer in Victor Hugo's The Hunchback of
Notre Dame.
Yogsothoth Shishkin & Zlatogursky,
2018 (centrohelid) "refers to Yog-Sothoth - the character of
the novels and short stories of Howard Phillips Lovecraft, which was
described as a conglomeration of glowing spheres." In the family
Yogsothothidae. [Protist 169: 684]
Ampulex dementor Ohl, 2014 (cockroach wasp)
Named after the Dementors of the Harry Potter books, which suck their
prey's souls much like the wasp's venom neutralizes its victim's
behavior.
Aname aragog Harvey et al., 2012 (trapdoor
spider) Named after a giant spider from Rowling's Harry Potter and
the Chamber of Secrets. [Zootaxa 3383:
15]
Lycosa aragogi Nadolny & Zamani, 2017
(wolf spider) Named after Aragog, from the "Harry Potter" series,
which was based on a wolf spider. [Zootaxa 4286:
597]
Ochyrocera aragogue Brescovit et al. 2018
(spider) "refers to Aragog, a spider capable of communicating with
humans and a lover of human flesh, from the literary classic 'Harry
Potter and the Chamber of Secrets', by J.K. Rowling."
Aquilonifer Briggs et al., 2016 (Silurian
arthropod) Its tethered appendages, interpreted as juveniles held in a
brooding behavior, inspired the name which translates literally as
"kite-bearer", a reference to Khalid Hosseini's novel The Kite
Runner.
Buratina Khramov, 2019 (Cretaceous
neuropteran) "Named after Buratina, a long-nosed character of the
science-fiction novel by Michael Kharitonov" [presumably The
Golden Key, or the Adventures of Buratina].
Calumma tarzan Gehring et al., 2010
(chameleon) So named partly because it was found in what was known
locally as the Tarzan Forest in Madagascar, and partly in hopes that
the name, like Tarzan himself, will promote conservation of the
species and the forest. [Salamandra 46: 167]
Cavisternum bom and
Pelicinus tumpy Ranasinghe & Benjamin,
2018 (spiders) Bom and Tumpy are goblins in "The Goblin
Looking-Glass" by Enid Blyton (1947).
Pelicinus snooky Ranasinghe & Benjamin,
2018 (spider) After the goblin and main character in "The
Firework Goblins" by Enid Blyton (1971).
Ischnothyreus chippy,
Silhouettella snippy and
S. tiggy Ranasinghe & Benjamin,
2018 (spider) Named for brownies in the story "Billy's Little
Boats" by Enid Blyton (1971). [Evolutionary Systematics 2:
65]
Cetiocaridae (extinct anomalocarid-like
arthropods) The book All Your Yesterdays by C. M. Kosemen
compiles artwork showing speculative but plausible ideas about extinct
animals. It included the "Ceticaris", a hypothetical
animalocarid-like suspension-feeder, drawn by John Mezsaros. The
Cetiocaridae is named after it.
Cthulhu James and Keeling, 2012
(parabasalid, a flagellated protist termite symbiont) Named for the
tentacled demon from the writings of H. P. Lovecraft. The protist's
flagellar bundle is reminiscent of the demon.
Nanocthulhu lovecrafti Buffington, 2012
(wasp) "Cthulhu's is described as having 'a pulpy, tentacled head,'
and the clypeal fuscina [on the front of the wasp's head] described
herein is reminiscent of Cthulhu's head"; "nano-" refers to the wasp's
small size; and H.P. Lovecraft, fiction author, created
Cthulhu. [Proc. Entomol. Soc. Wash. 114: 5]
Pimoa cthulhu Hormiga, 1994 (spider) Named after
H. P. Lovecraft's evil god.
Sollasina cthulhu Rahman et al., 2019
(Silurian echinozoan) Because tentacles.
Cthylla James and Keeling, 2012 (parabasalid, a
flagellated protist termite symbiont) In H. P. Lovecraft's writing,
Cthylla was the secret daughter of Cthulhu. The protist is a
smaller and simpler relative of the protist genus Cthulhu.
[PLOS One 8(3) (2013)]
Ochyrocera atlachnacha Brescovit et al. 2018
(spider) "refers to Atlach-Nacha, a supernatural entity from Cthulhu
mythology that resembles a huge spider with an almost human
face."
Dracorex hogwartsia Bakker et al. 2006
(pachycephalosaur dinosaur) Named for Hogwarts School of Harry Potter
fame. The genus means "dragon king." J. K. Rowling wrote, "I am
absolutely thrilled to think that Hogwarts has made a small (claw?) mark
upon the fascinating world of dinosaurs." The skull is on display at
the Children's Museum of Indianapolis.
Eeyorius Paulin, 1986 (Australian fish) named
for the donkey in A.A. Milne's Winnie the Pooh books.
Like Eeyore, it lives in damp, dark places.
Eriovixia gryffindori Ahmed et al. 2016
(spider) Because the spider resembles the Sorting Hat from Rowling's
Harry Potter books, it is named after Godric Gryffindor, the hat's
original owner. "An ode from the authors, for magic lost, and
found, in an effort to draw attention to the fascinating, but oft
overlooked world of invertebrates, and their secret
lives." [Indian J. Arach. 5: 25]
Irritator challengeri Martill, Cruikshank, Frey, Small &
Clarke, 1996 (small theropod dinosaur) "challengeri" refers to
Professor Challenger, a character from Doyle's The Lost World.
The generic name came from the fact that inept Brazilian fossil
collectors broke the skull in extracting it and rebuilt it
incorrectly: "From irritation, the feeling the authors felt
(understated here) when discovering that the snout had been
artificially elongated".
Ituglanis macunaima Datovo & Landim, 2005
(catfish) "From the modernist Brazilian masterpiece by Mário de
Andrade -- 'Macunaíma: o herói sem nenhum
caráter' -- meaning the hero without any character, in
reference of the absence of any exclusive (taxonomic) character for
the new species. Mário de Andrade's Macunaíma was based
in folk Amazonian indian myth, and also presents infantile features,
in allusion to the paedomorphic characters of the new
species." [Neotrop. Ichthyol. 3: 461]
Laelius arryni, L. baratheoni, L. lannisteri,
L. martelli, L. targaryeni, L. tullyi, and
L. starki Azevedo & Barbosa, 2014
(bethylid wasps) "The specific epithets of all new species are derived
from some families of the book A Song of Ice and Fire, Game of
Thrones [by George R. R. Martin]". [Zoologica (Curitiba)
31(3)]
Eadya daenerys Ridenbaugh, 2018 (wasp)
Named for Daenerys Stormborn of House Targaryen, from George
R.R. Martin's books and the Game of Thrones TV
series.
Pheidole drogon and
Pheidole viserion Sarnat, Fischer & Economo,
2016 (ant) Named for dragons (one black, one cream and gold)
of Daenerys Targaryen, a fictional character from the George
R. R. Martin's novel A Song of Ice and Fire. [PLoS ONE,
11:e0156709]
Gymnetis drogoni, G. rhaegali and
G. viserioni Ratcliffe, 2019 (scarabs)
Named after "Game of Thrones" dragons because the beetles' orange
markings suggested fire.
Ochyrocera varys Brescovit et al. 2018
(spider) "refers to Varys, a fictional character in George
R. R. Martin's book, 'A Song of Ice and Fire'. Lorde Varys is a
character with a venomous spirit, known as a spider in the
plot."
Paramonovius nightking Li & Yeates,
2018 (bee fly) "This species is named after the Night King in
the American fantasy drama Game of Thrones, because all the
specimens were collected in winter and the fly is mostly covered in
thick pale pruinescence." Also, the fly, like the fictional
villain, turns its victims into zombies. [Austral Entomo. 58:
192.]
Tritonia khaleesi Silva et al. 2013 (sea
slug) Its silvery back stripe suggested to the authors the platinum
blonde hair style of Khaleesi Daenerys Targaryen in the fantasy novel
series A Song of Ice and Fire, as played by Emilia Clarke in
the TV adaptation Game of Thrones.
Ledermanniella maturiniana Beentje, 2005 (a
minute Kenyan waterweed, Podostemaceae) Named after Patrick O'Brian's
character Doctor Stephen Maturin, an avid naturalist and pathetic sailor
who often managed to fall off boats. Like him, this plant is often
immersed.
Loraxichthys Salcedo, 2013 (catfish)
Named for Lorax, the Dr. Seuss character who advocates for the
environment; "Loraxichthys refers to the fish that speaks for
other fishes." [Zootaxa 3640: 565]
Macrocarpaea apparata Grant & Struwe,
2003 (gentian) Named after the verb "to apparate" made popular in
J.K. Rowling's Harry Potter series. As a wizard apparating out of
nowhere, this 12-foot high plant appeared to botanists on a misty
hillside in southern Ecuador. (More info
here)
[
Harvard Papers in Botany 8: 61]
Clevosaurus sectumsemper Klein et al.,
2015 (triassic lizard) Named for the "sectumsempra" spell from
Harry Potter novels, which causes an invisible sword to slash the
victim, much as the reptile would slash with its sharp teeth.
(Clevosaurus is named for Clevum, the Latin name of
Gloucester.)
Ochyrocera charlotte Brescovit et al. 2018
(spider) "refers to Charlotte, the spider from the classic
'Charlotte's Web' by E.B. White and a great friend of the pig named
Wilbur."
Ochyrocera misspider Brescovit et al. 2018
(spider) "refers to Little Miss Spider, a very popular spider around
the world and the main character of the children's books by David
Kirk."
See also O. aragogue, O. atlachnacha, O. varys,
O. larcana, and O. ungoliant elsewhere in
this section. [ZooKeys 726: 87]
Oileus gasparilomi Cano & Schuster,
2012 (bess beetle) "Named after Gaspar Ilóm, a native
hero of the novel 'Men of Maize' by [literature Nobel prize-winner]
Miguel Ángel Asturias. The collection locality is called
'mountains of Ilóm'." [Zookeys 194: 81]
Ozraptor subotaii Long & Molnar, 1998
(theropod dinosaur) Named after Subotai, a swift-running thief from
Conan the Barbarian, whose behavior this dinosaur is supposed to have
emulated. ("Oz" in the genus name comes from a colloquial abbreviation
of Australia.)
Potamalpheops tigger Yeo & Ng, 1997
(shrimp) Named for Tigger, the tiger character in
A.A. Milne's Winnie the Pooh books, because of "the bold
striped appearance of freshly caught live specimens."
[J. Nat. Hist. 31: 186.]
Savignia naniplopi Bosselaers and Henderickx
2002 (linyphiid spider) "The species is named after the gnome
(Latin 'nanus') Plop, a popular character from children's stories
whose cap is similar in shape to the male cephalic snout of the
present species." [Zootaxa 109:3]
Theognete maturini Anderson, 2010 (weevil)
Named for Stephen Maturin of Patrick O'Brian's Aubrey-Maturin series
of British Royal Naval novels.
Tinkerbella nana Huber & Noyes 2013
(fairyfly) Named for characters from J. M. Barrie's "Peter Pan":
the fairy Tinkerbell, and Nana, the nurse.
Neolebouria tinkerbellae Thompson &
Margolis, 1987 (microscopic trematode)
Pseudolucia hazeorum Bálint and Johnson,
1993 (lycaenid) Referring both to the hazy wing color and to the
Haze family from
Lolita.
Nabokov himself was an expert on lycaenids, particularly the genus
Pseudolucia, which he named. Other
lycaenid
names derive from Nabokov stories, too, including:
Madeleinea nodo, M. odon Bálint & Johnson,
1994 (lycaenid butterflies) Named after half-brothers in Vladimir
Nabokov's Pale Fire, reflecting their close
relatedness.
Madeleinea cobaltana Bálint & Lamas,
1994 is named for Kobalt, a mountain resort in Pale
Fire.
Madeleinea lolita Bálint, 1993,
Pseudolucia charlotte, P. clarea Bálint
and Johnson, 1993,
P. humbert Bálint and Johnson, 1995
(lycaenids) Named after characters in Nabokov's
Lolita.
Humbert humberti Sime & Wahl, 2002
(ichneumonid wasp) Named for Nabokov's Lolita pederast Humbert
Humbert. [Zool. J. Linn. Soc. 134: 1]
Paralycaeides hazelea Bálint & Johnson,
1995 and
P. shade Bálint, 1993 (lycaenids)
After characters in Nabokov's Pale Fire.
Itylos pnin Bálint, 1993
for Professor Pnin.
Nabokovia ada Bálint & Lamas,
1994, for the title character, and
Madeleinea ardisensis Bálint & Lamas,
1996 named after Ardis Hall, a place in Ada or Ardor: A
Family Chronicle.
Thestral Faúndex & Rider, 2014
(pentatomid bug) Named for the thestrals, skeletal winged horses, from
J. K. Rowling's Harry Potter series. The ivory carinae and calluses
on the bug's back resemble the skeletal body. Also, thestrals cannot
be seen by everyone, and few specimens of this insect have been found,
even in well-sampled locations. [Zootaxa 3884: 395]
Trimeresurus salazar Mirza et al. 2020
(pit viper) Named for Salazar Slytherin, a co-founder of Hogwarts
School of Witchcraft and Wizardry (in Rowling's Harry Potter series)
whose Parselmouth ability linked him to serpents.
Aleiodes adorabelleae,
A. anguaae,
A. atuin,
A. binkyi,
A. deathi,
A. lipwigi,
A. magratae,
A. morti,
A. ponderi,
A. pteppicymoni,
A. ridcullyi,
A. rincewindi,
A. stibbonsi,
A. tmaliaae,
A. vetinarii Butcher et al. 2012
(braconid wasps) - all names from Terry Pratchett's Discworld series.
[Zootaxa 3457]
Czekanowskia anguae Watson et al., 2001
(fossil gymnosperm) for Angua, she-werewolf and member of the
Ankh-Morpork City Watch. "In tribute to the author Terry Pratchett
OBE, all the new fossil plant species diagnosed and described in this
paper are named for fictional characters who appear in his series of
Discworld novels." [Bull. Nat. Hist. Mus. Lond. (Geol.) 57:
29-82.] (This refers also to the eight entries which
follow.)
Ginkgoites weatherwaxiae Watson et al.,
2001 (fossil ginkgo) for Granny Weatherwax, a
witch.
Ginkgoites nannyoggiae Watson et al., 2001
(fossil ginkgo) for Nanny Ogg, matriarch and witch.
Ginkgoites garlickianus Watson et al., 2001
(fossil ginkgo) for Magrat Garlick, witch and Queen of
Lancre.
Phoenicopsis rincewindii Watson et al.,
2001 (fossil gymnosperm) for Rincewind, an ineffective wizard of
Unseen University.
Pseudotorellia vimesiana Watson et al.,
2001 (fossil conifer) after Sir Samuel Vimes, commander of the
Ankh-Morpork City Watch.
Sciadopityoides greeboana Watson et al.,
2001 (fossil gymnosperm) After Greebo, Nanny Ogg's
cat.
Sulcatocladus dibbleri Watson et al., 2001
(fossil conifer) for C.M.O.T. Dibbler, street vendor in
Ankh-Morpork.
Torreyites detriti Watson et al., 2001
(fossil conifer) for Detritus, troll and member of the Ankh-Morpork
City Watch.
Adelopsis dumbo Gnaspini & Peck 2001
(leiodid beetle) Named for the big-eared cartoon elephant, because the
beetle's aedeagus, which resembles an elephant proboscis, has at its tip
a very large lateral projection resembling an ear.
Siphopteron dumbo Ong & Gosliner, 2017
(bat-winged slug) "refers to the similarity of this species to the
Disney character, Dumbo the elephant, as it swims through the water."
[Zool. J. Linn. Soc. 180: 755,
doi.org/10.1093/zoolinnean/zlw018]
Agra lilu Erwin, 2000 (ground beetle) Named
for a character in the film "The Fifth Element".
Anelosimus biglebowski Agnarsson, 2006
(spider) "after the movie 'The Big Lebowski' of Joel and Ethan
Coen." [Zootaxa 1147: 1]
Anelosimus dude Agnarsson, 2006 (spider)
"after 'The Dude', a character in the movie 'The Big
Lebowski'"
Aptostichus sarlacc Bond 2012 (trapdoor
spider) The sarlacc is a creature from "Star Wars Episode VI: Return
of the Jedi" which lives at the base of a sand pit, consuming people
and animals that fall (or are thrown) in.
Axima sidi Arias-Penna et al., 2014
(stalk-eyed wasp) Named after Sid, a bulbous-eyed sloth from the "Ice
Age" movies, because of his facial resemblance to the wasp.
[Zootaxa 3866: 588]
Bambiraptor Burnham, Derstler, Currie, Bakker, Zhou & Ostrom,
2000 (theropod dinosaur) after Disney's Bambi, because of its
small size. [
U. Kansas Paleo contributions 13] (See also a
dinosaur
mailing list thread which includes much discussion of the
appropriateness of the name.)
Begonia amidalae Lin et al. (begonia)
Named after Padme Amidala. Silvery spots on the plant's dark leaves
evoked a starry sky and inspired the name.
Xenokeryx amidalae Sánchez et al.,
2015 (Miocene ruminant) Named after Queen Padme Amidala from
"Star Wars: The Phantom Menace" because its horn (the genus name means
"strange horn") resembles one of her hairstyles.
Cambroraster falcatus Moysiuk & Caron,
2019 (Cambrian radiodont) Named for its resemblance to
the Millennium Falcon from the "Star Wars"
franchise.
Celmus michaelmus Adrain & Fortey, 1997
(trilobite) Its abdominal apex looks like a Mouseketeer
hat.
Ceraeochrysa michaelmuris Adams & Penny (lacewing)
Its abdominal apex looks like a Mouseketeer hat.
Chloridops regiskongi James & Olson, 1991 (extinct
Hawaiian finch) Described by a local journalist as "a real King Kong
finch", thus the name.
Cystomastacoides kiddo Quicke & Butcher, 2013
(braconid wasp) Named after the assassin character Beatrix Kiddo in
Quentin Tarantino's "Kill Bill" films. [J. Hymenop. Res. 31:
65]
Darthvaderum Hunt, 1996 (oribatid mite)
"Etymology: When I saw the SEM [scanning electron micrograph] of the
gnathosoma I immediately thought of Darth Vader, evil antihero of Star
Wars." [Records of the Australian Museum 48:
303-324]
Agathidium vaderi Miller and Wheeler, 2004
(slime mold beetle) Its head resembles Darth Vader's
helmet.
Begonia darthvaderiana Lin et al.
(begonia) It has leaves that are nearly black.
Polemistus vaderi Menke, 1983 (wasp)
Ricinus vaderi Valan, 2016 (louse)
[Parasite 23: 6]
Diplacodon gigan Mihlbachler, 2011
(brontothere) From the etymology: "'Gigan' is a fictional giant horned
monster first appearing in the 1972 Japanese film 'Godzilla versus
Gigan' and other Godzilla films thereafter." [J. Vert. Paleo.
31: 202]
Eoperipatus totoro Oliveira et al., 2013
(velvet worm) "the species is named after the main character of the
cartoon movie 'My Neighbour Totoro' by Hayao Miyazaki (1988, studio
Ghibli), who uses a many-legged animal as a vehicle..."
[Zoologischer Anzeiger 252: 495]
Godzillius Yager, 1986 (remipede crustacean) These are
the largest such crustaceans, from underwater caves in the Bahamas. The
family Godzilliidae takes its name from this genus.
Godzilliognomus Yager, 1989 (godzilliid) The
smallest remipede. [Bull. of Marine Sci.
44(3):1195]
Gojirasaurus Carpenter, 1997 (theropod
dinosaur) "Gojira" is the Japanese name for Godzilla (but the dinosaur
was found in New Mexico).
Pleomothra Yager, 1989 (godzilliid) Named after
Mothra. [Bull. of Marine Sci. 44(3):1195]
Gamerabaena Lyson and Joyce, 2010 (late
Cretaceous turtle) Named for the Gamera, the giant Japanese film
monster, and the family Baenidae of which it is a
member.
Microgaster godzilla Fernandez-Triana et
al. 2020 (parasitoid wasp) Named after Godzilla mainly because
of the monster being an icon of Japanese culture (the wasp is from
Japan); the wasp's hunting behavior also bears loose resemblance to
Godzilla's emergence from water and his relationship with
Mothra.
Sinemys gamera Brinkman & Peng, 1993 (Japanese
fossil turtle) Named after the giant Japanese fire-breathing flying
turtle. The fossil has wing-like projections from its shell.
Han solo Turvey 2005 (agnostid trilobite)
Officially, the genus is named after the Han Chinese (the fossil is from
northern Hunan Province, China), and the species is so named because it
appears to represent the last surviving member of the Diplagnostidae.
Really, Turvey's friends dared him to name a species after a Star Wars
character, as most of the characters' names sound like scientific names.
[Trans. Royal Soc. Edinburgh: Earth Sci. 95:
527-542]
Hirudicryptus quintumelementum Korsós et
al. 2008 (millipede) As the fifth named species of the genus,
it is named after the movie "The Fifth Element" ("Le
cinquième élément", 1997). The head and
collum of the new species also superficially resemble the alien
custodians of the movie. [J. Nat. Hist. 43: 435]
Hortipes terminator Bosselaers & Jacque
(spider) The male's palps resemble a "futuristic gun."
Ikrandraco avatar Wang et al., 2014
(Cretaceous pterosaur) Named for the Ikran, a flying creature from the
movie Avatar which shows a well-developed dentary crest similar to the
pterosaur's (+ draco, "dragon"). [Scientific Reports
4, no. 6329]
Maratus nemo Schubert, 2021 (peacock
spider) So called because the male's coloring resembles that of the
title character of the film "Finding Nemo".
Osedax jabba Rouse et al., 2018 (bone worm)
"The trunk of the new species is reminiscent of the tail of the
mythical creature Jabba the Hutt from the Star Wars
franchise." [Zootaxa 4377: 476]
Peckoltia greedoi Armbruster, Werneke &
M. Tan, 2015 (catfish) Named after Star Wars bounty hunter
Greedo, whom it resembles. [ZooKeys 480: 97]
Polemistus chewbacca Menke, 1983
(wasp)
Trigonopterus chewbacca Van Dam & Riedel,
2016 (weevil) Its dense scales remind the authors of the
Wookie's dense fur. [ZooKeys 582: 129]
Wockia chewbacca Adamski, 2009 (moth)
Predatoroonops Brescovit et al., 2012
(goblin spider) The spiders' chelicerae resemble the face of the
Predator from the 1987 film of that name, hence the genus name. The
17 species described within the genus are all named after characters,
actors, locations, and other names from the movie.
Prolatcyclus kindzadza Mychko et al. 2019
(Carboniferous cyclid crustacean) "In honor of the cult sci-fi
dystopian tragicomedy Soviet film 'Kin-dza-dza!' directed by
Georgiy Daneliya in 1986."
[N. Jb. Geol. Paläont. Abh. 294: 88.]
Qrocodiledundee outbackense Fernandez-Triana
& Boudreault (wasp) The name refers to the remote region of
Australia where the wasp is found and to Crocodile Dundee, a movie
character associated with such a region.
Ramisyllis kingghidorahi Aguado et
al. 2022 (branching annelid) King Ghidorah was a three-headed,
two-tailed monster enemy of Godzilla. Like the worm, it is from
Japan and can regenerate its lost ends.
Shrekin Britto and Navia, 2007 (eriophyid
mite) Named for the film cartoon character Shrek "because of the
resemblance of the long, laterodorsal scapular tubercules to the long
stalked ears of this character, plus -in, Latin diminutive
suffix." [Internat. J. Acarol. 33:347-351]
Stormtropis Perafán et al., 2018 (spider)
"... a Latin declension of the noun Stormtrooper from the fictional
universe of the Star War films. . . . These soldiers are very
similar to each other, with some capacity for camouflage but with
unskillful movements, like this group of spiders."
[ZooKeys 830:1]
Tetramorium jedi Hita Garcia & Fisher,
2012 (ant) Named for the Jedi of "Star Wars" fame.
[Zootaxa 3592: 51]
Xenomorphia Krogmann et al. 2018 (fossil
wasp) The wasps were discovered inside fossil fly pupae, having
eaten their hosts from the inside out, a behavior similar to the
Xenomorph alien of the Alien movie.
Dolichogenidea xenomorph Fagan-Jeffries & Austin,
2018 (wasp) The wasp lays eggs in a caterpillar, for its
larvae to each the caterpillar from the inside out. [J. of
Hymen. Res. 64:177]
Yoda Priede et al., 2012 (acorn worm) Named
for Yoda of Star Wars fame because large lips at the worm's head
resemble Yoda's ears. [Invert. Biol. 131:
244.]
Albunione yoda Markham & Boyko, 2003 (parasitic
isopod (Crustacea: Isopoda: Bopyroidea)) This species, found under the
carapace of sand crabs, also has large projectiog lateral flaps on the
sides of its head that look like Yoda's ears.
Polemistus yoda Vincent, 1983
(wasp)
Trigonopterus yoda Riedel, 2019 (weevil)
A small, greenish, forest-dwelling weevil.
[ZooKeys 828:1]
Zuul crurivastator Arbour and Evans, 2017
(Cretaceous ankylosaurid dinosaur) Its head resembles Zuul, Gatekeeper
of Gozer, a monster from the 1984 Ghostbusters. (The specific
epithet means "destroyer of shins". Arbour said, "I had really wanted
to use that for a long time, and I was saving it for a specimen with a
really well preserved tail.") [Royal Society Open Science
161086]
Agra dax Erwin, 2000 (ground beetle) Named
for the character Jadzia Dax from "Star Trek: Deep Space
Nine" and dedicated to the actress, Terry Farrell.
Agra smurf Erwin, 2000 (ground beetle) The
name was inspired by the beetle's head shape.
Annuntidiogenes worfi Fraaije et al., 2009
(Cretaceous hermit crab) The ornament of this crab recalls the
forehead of Star Trek's Mr. Worf. [Bol. Soc. Geol. Mex.
61: 13]
Conus tribblei Walls, 1977 (marine snail) Named after
a pet cat named "Tribbles", which was named after the furry creatures
from Star Trek. [The Pariah 1: 1-3.]
Elthusa xena van der Wal, 2019 (fish
parasite isopod) "named after Xena, the warrior princess, in
reference to the strong nature of the female cymothoid isopod."
[ZooKeys 841: 1]
Eucteniza caprica Bond & Godwin, 2013
(trapdoor spider) "in reference to the humanoid cylon model Caprica
6, portrayed by Tricia Helfer in the remake of the science fiction
series Battlestar Galactica." [ZooKeys356: 31]
Geragnostus waldorfstatleri Turvey 2005
(trilobite) The pygidium (tail) looks just like the heads of Waldorf and
Statler of "The Muppet Show." [Trans. Royal Soc. Edinburgh: Earth
Sci. 95: 527-542]
Ladella spocki Viegas, Benaim & Absãlo,
2014 (mussel) for Spock of Star Trek fame. "The
overall shape of [the mussel's] valves resembles the shape of the
pointed ear of the Vulcans ..."
[Am. Malacological Bull. 32: 188-189.]
Mestoronema Wagner 1999 (fossil snail) Named after the
evil snail king on a Dr. Who episode. [Smithsonian Contrib. to
Paleobiology 88:1-154] (Turnabout is fair play; the writers for
Dr. Who often took animal names for their monsters -- crinoids,
mandrills, Mara, etc.)
Choeras zygon Fagan-Jeffries et al., 2019
(braconid wasp) Named after the Zygon race of aliens from Dr. Who
because the Zygons, like the wasp larvae, consume their hosts while
inhabiting them.
Cyclocardia dalek Pérez & del
Río, 2017 (Cenozoic bivalve) Named after main villains
of Doctor Who, "characterized by an armour with prominent circles,
similar to the nodual external sculpture of this
species."
Phanuromyia odo Nesheim, 2017 (wasp)
Because of this species' variable morphology, it was named for a
shape-shifting character from Star Trek: Deep Space Nine.
[ZooKeys 663: 71]
Spongiforma squarepantsii Desjardin, Peay &
T.D. Bruns, 2011 (mushroom) This orange mushroom from Borneo was
so unusual in its spongelike form, the researchers named it after
Spongebob Squarepants, the world's most famous sponge. Additionally,
its spore-producing area, under high magnification, resembles the
seafloor where Spongebob lives.
Astrolirus patricki Zhang et al. 2020
(starfish) All specimens were found on sponges; the species is
named after the character Patrick Star, who always spends time with
his friend SpongeBob Squarepants.
Yada Yada (alphavirus) The virus infects only
mosquitoes. Named after a catchphrase from Seinfeld, its
discoverers say it just isn't very exciting (but, one adds,
"Seinfeld is awesome").
Adetomyrma goblin Yoshimura & Fisher,
2012 (ant) Adults of this blind subterranean ant drink the
hemolymph of their own larvae. Only the larvae can eat solid
food.
Advhena Castello-Branco et al. 2020
(sponge) From Latin advena, "alien", because the sponge's
shape resembles a popular Hollywood depiction of aliens. (The
name Advena was already taken by a gastropod.)
Agra sasquatch Erwin, 1982 (carabid) with big
feet.
Agra yeti Erwin, 1982 (carabid) sister
species of A. sasquatch.
Mesocentrus sasquatch Butcher et al. 2014
(braconid wasp) from Papua New Guinea. It was not explained why that
name was chosen. [Zootaxa 3860: 449]
Boana icamiaba Peloso et al., 2018
(gladiator frog) Named for the Icamiabas, a legendary female-only
warrior tribe from Brazil. [S. Am. J. of Herp.
13(2)]
Cryptocellus icamiabas Tourinho & de
Azevedo, 2007 (mite) Named for the Icamiabas, female warriors
described in the chronicles of the Dominican friar Gaspar de Carvajal,
which has been identified as source of the name of the Amazon River
and the Brazilian state of Amazonas. The mite was discovered in
Amazonas. [Zootaxa 1540: 56.]
Camelotia Galton, 1985 (Triassic prosauropod)
from England; named for Camelot.
Campsicnemius uncleremus Evenhuis, 2000
(dolichopodid fly)
Chupacabrachelys complexus Lehman & Wick,
2010 (Cretaceous turtle) The chupacabra (Spanish
for "goat sucker") is a creature of contemporary folk legend said to
feed on livestock in the Texas-Mexico border area. The skull of
Chupacabrachelys, to the authors, resembles a mangy coyote
believed to be responsible for chupacabra sightings. The
complexus epithet refers to "The Complex" tour of the Blue Man
Group, which entertained the authors during their work.
Cibotium barometz (L.) J.Sm. (woolly fern)
The barometz, or Vegetable Lamb of Tartary, was an imagined plant
whose fruit grew into sheep which, while connected to the plant by an
umbilical cord, grazed the area around it. Rhizomes growing up from a
woolly common base of the actual fern can form the shape of an
inverted lamb.
Cindarella Chen et al., 1996 (Cambrian
arthropod) Named "for its phonetic similarity to Xandarella" (a
similar Cambrian fossil), but likely influenced by the name of the
folkloric woman. [Ramsköld et al. 1997, Trans. Royal
Soc. Edinburgh Earth Sci. 88: 25.] Xandarella itself is
the diminutive of Greek Xandaros, "a fabulous sea-monster" [Hou et
al., Zool. Script. 20: 402].
Cinderella Steyskal, 1949 (heleomyzid
fly)
Crocidura cinderella (Cinderella shrew) from
Africa.
Teleocichla cinderella Kullander 1988
(cichlid) "Named with reference to the coloration, gray and black,
producing an ashy appearance" [Copeia,
196-230].
Thylamys cinderella (Cinderella fat-tailed
opossum) from NW Argentina.
Cyrtodactylus rukhadeva Grismer et
al. 2021 (gecko) The Rukha Deva, in Thai folklore, are sylvan
spirits which live in or on old trees.
Dracaena draco (dragon tree)
Dracaena and
Draco (lizards)
Dracunculus (round worm) Named after "draco", dragon.
D. medinensis is the largest tissue parasite of man (it can grow
longer than 3 feet). It is possible to extract the worm by winding it
slowly, over a period of days or weeks, around a stick. This may be the
source of the physician's caduceus. D. medinensis is on WHO's hit
list and may soon be eradicated.
Dragonana (leafhopper)
Kryptodrakon Andres, Clark & Xu, 2014
(Jurassic pterosaur) Translation: "Hidden dragon"; so named because it
was found in the area where "Crouching Tiger, Hidden Dragon" was
filmed.
Dendrobium nagataksaka Metusala (orchid)
"... 'naga' means dragon, and 'taksaka' is the name of the
ancient Indian, Javanese and Balinese mythical dragon. It refers to
the shape of the flower which has erect petals and long protruding
lip resembling the dragon’s head with long horns and long
lip."
Qilianglong Xing et al., 2015 (Jurassic
Sauropod) meaning "dragon from Qijiang".
Eviulisoma nessiteras Enghoff, 2018
(millipede) "Nessiteras is the genus name given to the famous
Loch Ness Monster, and the dorsal lobe of the solenophore
of E. nessiteras sp. nov. resembles the most famous
photograph of the alleged monster sticking its long neck out from
the lake surface." [Eur J Taxon. 445:40]
Gargoyleosaurus Carpenter, Miles, & Cloward,
1998 (ankylosaurid dinosaur)
Goniacodon hiawathae Van Valen, 1978
(paleocene mammal) for Hiawatha, legendary founder of the Iriquois
League.
Kikimora palustris Eskov, 1988 (spider)
Kikimora is a dangerous female spirit in Slavic mythology who lives in
marshes. "Palustris" means "of a marsh." [Zoologicheskyj
Zhurnal 67: 678 (in Russian)]
Lathrobium hibagon Senda, 2020 (rove
beetle) Named after the Hibagon, a Japanese equivalent of a Bigfoot
or yeti. Sightings, which began in the 1970s, occur mainly in the
Mt. Hiba-yama area, the type locality of this species.
[Jap. J. of Syst. Entomo. 26: 186.]
Leprechaunus (treehopper)
Merlinia (trilobite) Named for King Arthur's
wizard
Metapheretima durendali and
M. excaliberi Easton, 1979 (earthworms)
Durendal was the sword of Roland; Excalibur, of King Arthur. (See
more sword eponyms in the Tolkien section below.)
Excalibosaurus McGowan, 1986 (Jurassic ichthyosaur)
Named after King Arthur's sword. Excalibosaurus has a swordlike
upper jaw, and it was found in Britain's west country, the place of the
emergence of Excalibur.
Lemur Linnaeus, 1758 (lemur) From
Latin leumures, zombie-like ghosts which the Romans supposed
ultimately would haunt the world at night. The name was first applied
to the slender lorises of India, inspired by their slow movement and
nocturnal habits. Its meaning later narrowed to just the Malagasy
primates.
Lusitanipus xanin Gilgado, 2020
(millipede) A Xanín is a goblin-like creature said to inhabit
the Spanish forests where the millipede was found. Like the
millipede, the Xanín is small, lives in the shade, and tries
to hide.
Monstera Adans. (tropical vine) A
cultivated species with edible fruit has the ironic name
Monstera deliciosa.
Mucha tzokotucha Ozerov, 1992 (fly:
Sepsidae) Named for a fly character in a famous Russian folktale.
"Mucha" means "fly", and "tzokotucha" is the character's nickname for
himself; apparently it has no further meaning.
Paroxyna babayaga Hering, 1938 (tephritid
fruit fly) Named for the ugly Russian folklore monster
Babayaga.
Polypterus mokelembembe Schafer and Schliewen
2006 (freshwater fish called bichir or reedfish) Named for the
cryptozoological Congolese dinosaur-like creature Mokele-mbembe
(featured, e.g., in the Disney movie "Baby"). The bichirs have been
around since the time of the dinosaurs, and this particular species
comes from the Congo. [Zootaxa 1129: 25-36]
Prosopanche demogorgoni Funez (forb) So
called because its flower resembles the fictional demogorgon
monster, presumably as depicted on Netflix's "Stranger Things,"
although that monster is usually depicted with five sepals, and the
flower has three.
Protomelas krampus Dierickx & Snoeks,
2020 (fish) Named for the European folklore character Krampus,
who puts naughty children in a bag and takes them away, because this
fish feeds on eggs and larvae of other fish. The goat-like
appearance of Krampus also evokes to the fish's butting
behavior.
Samrukia Naish et al. 2011 (Cretaceous
bird) Named for the Samruk, a magical bird of Kazakh
folklore.
Sanctacaris Briggs & Collins, 1988 (fossil primitive
chelicerate) Literally "Santa claws" [see Gould, Wonderful Life,
p. 186-187]
Selenochlamys ysbryda Rowson & Symondson, 2008
(ghost slug from Wales). The species name is Latinised from the
Welsh ysbryd (meaning ghost or spirit), referring to the fact
that it is rarely seen, is white in color, and is
nocturnal.
Smok wawelski Niedzwiedzki, Sulej and Dzik,
2011 (Triassic archosaur) Named for Smok wawelski (the dragon of
Wawel Hill), a famous dragon in Polish folklore.
Tarascosaurus salluvicus Le Loeuff &
Buffetaut, 1991 (Cretaceous theropod) Named for the Tarasque, a
medieval legendary dragon from Provence, France. The Salluvii were an
ancient Gallic tribe in the area near Marseilles.
Taraschelon Pérez-García,
2016 (Oligocene tortoise) Also named after the
Tarasque.
Tetramorium elf Hita Garcia & Fisher,
2012 (ant)
Thitarodes shambalaensis Wang et al. 2019
(moth) Named for Shambala of Tibetan tradition, the mythical kingdom
hidden in the mountains, referring to the species' alpine
habitat under Mt. Gongga.
Troglocladius hajdi Andersen et al., 2016
(midge) This midge, 1,431 meters deep in a cave in southern Croatia,
the only blind cave insect that flies, is named after the Hajdi,
winged cave-dwelling dwarf-like creatures from Slavic mythology.
[PLoS ONE 11(4)]
Tsoabichi Brochu, 2020 (Eocene
alligatorid) From Shoshone tso'abichi', "monster", in
reference to the species occurring in traditionally Shoshone
territory. [J. Vert. Paleo. 30: 1109.]
Vampyrodes (South American bat),
Vampyressa (South American bat),
Vampyrum (false vampire bat) The last
is also misnamed, for it does not feed on blood.
Vampyrellidae (protist)
Drepanomenia vampyrella (Heath, 1905)
(solenogaster mollusc)
Pteropus vampyrus (giant Malaysian fruit
bat)
Rhacophorus vampyrus (frog)
Vouivria Mannion et al. 2017 (Jurassic
brachiosaur) "The generic name is derived from the old French word
'vouivre', itself from the Latin 'vipera', meaning 'viper'. In
Franche-Comté, the region in which the holotype was
discovered, 'la vouivre' (=the wyvern) is a legendary winged
reptile. In the homonym novel written by the great French author
Marcel Aymé, 'La Vouivre' is a beautiful woman who lives in
the swamps in the neighbourhood of Dôle (Franche-Comté)
and protects a spectacular ruby."
(See also "Named after Writers")
Aletodon mellon (Van Valen, 1978)
(Paleocene mammal) "mellon," Elvish for "friend," was the password into
Moria.
Ancalagon Conway Morris, 1977 (Cambrian priapulid)
From a dragon from Tolkien.
Ankalagon Van Valen, 1980 (Paleocene
mesonychid mammal) Renamed from Ancalagon because it was
preoccupied.
Anisonchus eowynae Van Valen, 1978 (Paleocene
mammal, synonym of A. athelas Van Valen 1978) for Éowyn,
Princess of Rohan. "Athelas" was a Middle Earth healing
plant, used to cure Éowyn.
Aspidoras azaghal Tencatt et al. 2020
(catfish) Named after a First Age dwarf.
Aspidoras psammatides Britto et al. 2005
(fish) "Named after 'Psammatos psammatides' [Psamathos Psamathides],
the sand sorcerer, a characer of the J.R.R. Tolkien's
book Roverandom, from the Greek psammos, meaning sand,
and ides, meaning son of. In allusion to the sand-dwelling
behavior of the species.
Asthenodipsas lasgalenensis Loredo et
al. 2013 (Mirkwood forest slug snake) "The specific
epithet lasgalenensis is derived from the name Eryn
Lasgalen which means in the 'Wood of Greenleaves' in the
fictional Sindarian language from J.R.R Tolkien's The Lord of the
Rings (1955). It was the name used by the Wood Elves for the
Mirkwood Forest after its cleansing following the War of the Ring.
This name was chosen because Tolkien's (1955) description of this
forest showed great similarity to the cloudy, upland forests within
which this species is found.
Balinia Hedqvist, 1978 (chalcidoid wasp)
Named after Balin, a dwarf.
Balrogia Hedqvist, 1977 (pteromalid
wasp)
Anthracosuchus balrogus Hastings et
al. 2014 (Paleocene crocodyliform) Found in a coal mine
(Anthracosuchus means "coal crocodile"), this giant croc was
named after the Balrog, another fearsome being which lived in a
mine.
Semicytherura balrogi Brouwers, 1994
Beorn Cooper, 1964 (fossil tartigrade) Named after the
man/bear character Beorn from The Hobbit.
Beornia Hedqvist, 1975 (wasp)
Bofuria Hedqvist, 1978 (pteromalid wasp)
Named after Bofur, a dwarf from The Hobbit.
Bomburia Hedqvist, 1978 (pteromalid wasp) for
Bombur
Bomburodon (Van Valen, 1978) (Paleocene mammal)
formerly named Bomburia, but Hedqvist's wasp had
priority.
Borophagus orc Webb, 1969 (Pliocene
canid) (Formerly in genus Osteoborus)
Bubogonia bombadili and
Protoselene bombadili (Van Valen, 1978)
(Paleocene mammals) after Tom Bombadil, "a simple, powerful, and
very old being."
Claenodon mumak (Van Valen, 1978) (Paleocene
mammal) after Mûmak, the Middle Earth elephant-like creature,
referring to size.
Desmatoclaenus mearae (Van Valen, 1978)
(Paleocene condylarth) "Meara, any one of the great horses of Rohan
in The Lord of the Rings."
Durinia Hedqvist, 1978 (chalcidoid wasp)
Named after Durin, a dwarf.
Deltatherium durini Van Valen, 1978
(Paleocene mammal) Several notable Dwarves were named Durin.
Dvalinia Hedqvist, 1977 (pteromalid wasp)
Named after Dwalin, a dwarf from The Hobbit, or after Dvalinn,
a dwarf from Norse mythology.
Earendil undomiel Van Valen, 1978 (Paleocene mammal)
"Eärendil, [father of Elrond], who, (in the Silmarillion)
sailed with a silmaril to get the aid that defeated Morgoth", and
"Quenya (Elvish), undómiel, evening star, which Eärendil
with his silmaril became."
Elachista amrodella, E. aredhella, E. caranthirella,
E. curufinella, E. daeronella, E. diorella, E. finarfinella,
E. gildorella, E. indisella, E. maglorella, E. miriella,
E. turgonella (Kaila 1999) (moths) Named after elves from
Tolkien, respectively: Amrod (Amras' twin), Aredhel (The White Lady of
Gondolin), Caranthir, Curufin, Daeron (Chief loremaster of Doriath),
Dior (King of Doriath), Finarfin (Noldor King in Aman), Gildor Inglorion
(High-Elf of Eriador & Imladris), Indis, Maglor, Miriel, Turgon (Lord of
Nevrast, then Gondolin). Kaila mentions that Elves "one after other
sailed over the water to the West, and were later difficult to see with
human eyes," alluding to the studied moths, which are very inconspicuous
and have spread to Nearctic areas. [Acta Zool. Fennica
211]
Entia Hedqvist, 1974 (wasp) Named after the
Ents. But it is a junior synonym of Boucekastichus, hence the
latter is the accepted name.
Feanora De Clerck & Schockaert, 1995
(turbellarian flatworm) from Fëanor, a personage from
the Silmarillion.
Fimbrethil ambaronae Van Valen, 1978
(Paleocene mammal, synonym of Oxyacodon agapetillus (Cope 1884))
Fimbrethil was an Ent-maiden; Ambaróna was a name for the Ents'
forest.
Frodospira Wagner 1999 (Silurian gastropod)
A small genus named after a certain hobbit. [Am. Malacological
Bull. 15:1-31]
Macrostyphlus frodo Morrone, 1994 (Andean
weevil)
Psylla frodobagginsi Martoni & Armstrong,
2019 (psyllid) So named because it is smaller than most
psyllids (as hobbits are smaller than humans), and because it was
found in New Zealand where P. Jackson's "Lord of the Rings" movies
were mostly filmed.
Gandalfia Willems et al. 2005
(platyhelminth)
Macrostyphlus gandalf Morrone, 1994 (Andean
weevil) [This and M. frodo are from American Museum
Novitates 3104: 1-63.]
Gervasiella oakenshieldi Paladini &
Cavichioli, 2015 (spittlebug) for Thorin
Oakenshield.
Gildoria Hedqvist, 1974 (wasp)
Named after Gildor Inglorion, an elf.
Gimlia Hedqvist, 1978 (chalcidoid wasp)
Named after Gimli, a dwarf.
Nebela gimlii Singer & Lara, 2015
(amoeba) referring to the dwarf Gimli because of its small size and
stout shape. "In addition, it has been found abundantly in a
forest, and Gimli was unique among his kind to have been travelling
in the woods."
Gollum Compagno, 1973 (catshark)
Gollumia Riedel, 1988
(snail)
Gollumiella Hedqvist, 1978 (chalcidoid
wasp)
Gollumjapyx smeagol (dipluran
hexapod)
Aenigmachanna gollum Britz et al. 2019 (fish)
The first known subterranean species of its family.
Galaxias gollumoides (fresh-water fish)
Named after Gollum because it has large eyes and was found in a
swamp.
Goniurosaurus gollum Qi et al. 2020
(cave gecko)
Ingerophrynus gollum Grismer 2007 (Mayasian
toad) So named for its likeness to the fictional amphibious
character. [J. Herp. 41: 225]
Gwaihiria Nauman (diapriid wasp) Named for
Gwaihir, Lord of the Eagles.
Khamul gothmogi Gates, 2008 (eurytomid wasp)
Named after Gothmog, Lord of the Balrogs.
Helferella gothmogoides Williams & Weir,
1988 (beetle)
Khamul Gates, 2008 (eurytomid wasp) Named
after Khaml, the Shadow of the East, the only Nazgul specifically
named by J. R. R. Tolkien. [Zootaxa 1898:
25]
Legolasia Hedqvist, 1974 (wasp)
Named after the elf Legolas.
Liolaemus tulkas Quinteros et al. 2008
(lizard) "in the mythology of J.R.R. Tolkien, 'Tulkas' is one of
the ainur or powers that helped shape arda or middle earth. One of
the characteristics of Tulkas is that of running faster than any
other creature. Liolaemus tulkas is very fast in short
sprints."
Litaletes ondolinde Van Valen, 1978
(Paleocene mammal) for Ondolindë, an Elven city.
Lotharingius frodoi Mattioli, 1996
(coccolithophore) after Frodo.
Metapheretima kilii, M. dorii Easton,
1979 (earthworms) Presumably named after the dwarves Kili and
Dori from The Hobbit. (Easton provides no
etymologies.)
Metapheretima elrondi Easton, 1979
(earthworm) Presumably named after Elrond, Lord of
Rivendell.
Metapheretima anduril, M. stingi, M. orcrista, and
M. glamdringi Easton, 1979 (earthworms)
Named for Anduril, re-forged sword of Aragorn; Sting, elven knife of
Bilbo and later Frodo Baggins; Orcrist, elven sword of Thorin
Oakenshield, and Glamdring, elven sword of Gandalf. (See also two
more sword eponyms in the Folklore section.)
[Bull. Brit. Mus. Zoology series 35: 1]
Mimotricentes mirielae Van Valen, 1978
(Paleocene mammal, synonym of Loxolophus hyattianus (Cope, 1885))
after Míriel, an Elf.
Mimatuta morgoth Van Valen, 1978 (Paleocene
mammal) for the "dark enemy of the world." Reference is to the
Hell Creek Formation.
Mimatuta minuial Van Valen, 1978 (Paleocene
mammal) "minuial" is Elvish for dawn's twilight. Reference is to
the dawn of the Cenozoic.
Mithrandir Van Valen, 1978 (Paleocene mammal) one of
the names of the wise wizard Gandalf. "Reference is to the
subtleness of the differences between the subgenera."
Nazgulia Hedqvist, 1973 (chalcidoid wasp)
Named after the Nazgul.
Abavorana nazgul Quah et al. 2017
(frog)
Tetramorium nazgul and
T. smaug Hita Garcia & Fisher, 2012
(ants)
Niphredil radagasti Van Valen, 1978
(Paleocene mammal, now in genus Paleotomus) Niphredil is a small
Middle Earth flower. Radagast the Brown was a wizard.
Ochyrocera laracna Brescovit et al. 2018
(spider) "refers to Laracna, a giant and very old spider created by
J. R. R. Tolkien in the classic book 'The Lord of the
Rings'."
Ochyrocera ungoliant Brescovit et al. 2018
(spider) "refers to Ungoliant, an evil spider spirit created by
J. R. R. Tolkien in the book 'The Silmarillion'."
Odontonia bagginsi de Gier & Fransen,
2018 (shrimp) Like the hobbits, the shrimp has hairy
feet.
Oinia Hedqvist, 1978 (chalcidoid wasp)
Named after Óin, a dwarf.
Oxyprimus galadrielae Van Valen, 1978 (arctocyonid
Paleocene mammal) for elf Lady Galadriel.
Pseudophallus galadrielae Dallevo-Gomes et
al. 2020 (pipefish) Named after Galadriel, Queen Elf
from Lord of the Rings. "The elf ruler of Lothlórien
is bearer of the ring Nenya, also known as the ring of water. It is
used herein in reference to the additional bony rings diagnostic of
the new species and its association with freshwater
habitats." [Zootaxa 4859: 95]
Paradzickia morwen Blagoderov, 1998
(fungus gnat) After the character Morwen.
Paradzickia hador Blagoderov, 1998
(fungus gnat) After the character Hador.
Paragiopagurus hobbiti (Macpherson, 1983)
(hermit crab)
Peperomia hobbitoides T. Wendt, 2003
(piper plant) "Peperomia hobbitoides is a small and humble
plant that lives in an almost fairyland-like environment of wet
karst outcrops in rain forest, and it is strongly and faithfully
tied to this home substrate. Indeed, it spends perhaps the greater
part of the year in holes and depressions in the rock as a resting
tuber. It is edible, an attribute of high esteem among hobbits. And,
like the hobbits, its home is under threat by forces much larger
than itself, in this case forest clearing . . ." [Lundellia
6: 37]
Saurodocus hobbit Yerman & Krapp-Schickel,
2008 (amphipod)
Syconycteris hobbit Ziegler, 1982 (moss-forest blossom
bat) "The specific name is an appositive noun, alluding to certain
analogies between the newly described species and another seldom-seen
forest form, described by Tolkien (1937), one of whose chief
distinguishing characteristics was also a thickly haired
pes."
Pericompsus bilbo Erwin (carabid) for the title
character of The Hobbit. So called because "it was short, fat,
and had hairy feet."
Platymastus palantir Van Valen, 1978 (Paleocene
mammal) "... one of 7 globes made by Fëanor that gave
visions through spacetime. Reference is to the long duration of the
genus."
Protungulatum gorgun Van Valen, 1978
(Paleocene mammal) "gorgûn" is a term for Orcs.
Sauron Eskov, 1995 (spider) Though found
in the Saur Mountain range (Kazakhstan), the name is derived from
the Tolkien character.
Sauroniops Cau et al., 2012 (Cretaceous
theropod dinosaur) The name refers to the "Eye of Sauron"; the
dinosaur is known only from a single bone above the eye
socket.
Litoria sauroni Richards & Oliver,
2006 (frog) From Tolkien's Sauron, "in reference to the
striking red and black mottled eye of this
taxon."
Macropsis sauroni Hamilton, 1972
(leafhopper)
Semicytherura tauron Brouwers, 1994
"After Tauron, a character in J.R.R. Tolkien's adventures of
Middle Earth."
Shireplitis Fernandez-Triana &
Ward, 2013 (braconid wasp) Named after the Shire (and the
related genus Paroplitis); endemic to New Zealand. Species
within the genus are:
S. bilboi, S. frodoi, S. meriadoci,
S. peregrini, S. samwisei,
and S. tolkieni.
Smaug Stanley et al., 2011 (cordylid lizard) Smaug
is the name of the dragon in The Hobbit. "According to
Tolkien the name is derived from the Old German verb
smeugen -- to squeeze through a hole. Like the type
species, Smaug lived underground and was heavily armored.
Appropriately Tolkien was born in the Free State province, South
Africa, the core area of distribution of the type species."
[Mol. Phylo. Evo. 58: 53]
Liolaemus smaug Abdala et al. 2010
(lizard) "In Tolkien's mythology Smaug, the Golden, is the last of the
Middle Earth dragons. The name Liolaemus smaug is because
this new species exhibit a golden coloration on body."
[Cuad. herpetol. 24]
Planois smaug Carvajal, Faúndez, and
Rider, 2015 (shield bug) Named after the dragon because of its
large size, because it had been "sleeping" in collections for about
60 years, and because its natural habitat resembles Middle
Earth.
Pycnophyes smaug Sánchez et
al. 2013 (kinorhynch)
Smeagol Climo, 1980 (gastropod, family Smeagolidae)
Smeagol was Gollum's original name.
Smeagolia Hedqvist, 1973 (pteromalid
wasp)
Iandumoema smeagol Pinto-da-Rocha et
al. 2015 (cave harvestman) One cave-dweller named after
another.
Thangorodrim thalion Van Valen (Paleocene mammal,
synonym of Oxyclaenus Cope 1884) "Thangorodrim, the
mountainous triple fortress of Morgoth in The Silrnarillion.
Reference is to Purgatory Hill [where the fossil was collected]."
Specific epithet "Sindarin (Elvish) thalion, strong.
Reference is to the massive morphology and the generic
name."
Tinuviel Van Valen (Paleocene mammal) for a most
beautiful elf. The name is Elvish for nightingale.
Yavanna Vera, 2013 (Cretaceous tree fern)
In Tolkien's Silmarillion, Yavanna is a godlike Ainur
who created the plants and animals of Middle Earth.
Batman Whitley, 1956 (Australian fish) Refers to a
similarity between this fish's dorsal fin and the "bats" with which
servicemen used to signal approaching aircraft on a carrier, rather than
the famous Dark Knight of the graphic novels. Batman has now been
replaced with the older Cryptocentrus.
Otocinclus batmani Lehmann, 2006 (catfish) is
named after the caped crusader, because of a bat-shaped mark on its
tail. [Neotropical Ichthyology 4: 379]
Campsicnemus popeye Evenhuis, 2013 (fly)
So named because its enlarged tibia resemble the bulging forearms of
the cartoon character Popeye.
Cirrhilabrus wakanda Tea et al. 2019
(vibranium fairy wrasse) Named for the fictional home of the
superhero Black Panther, which remained hidden from the world for a
long time. The common name "refers to the fictional metal
vibranium, a rare substance found on Wakanda that is woven into
Black Panther's suit. The purple chain-link scale pattern of the new
species is reminiscent of this detail." [ZooKeys 863:
85-96]
Cremnops wyleycoyotius Tucker et al. 2015
(wasp) Named after J. Wiley, who collected the specimen, and the
cartoon character Wile E. Coyote because it "sneakily (like the
well-known canine) remained undescribed until now."
Daptolestes bronteflavus, D. feminategus,
D. illusiolautus Robinson & Yeates, 2020, and
Humorolethalis Robinson, Li & Yeates,
2020 (robber flies) All have Marvel universe names. Thor's
fly,
D. bronteflavus, means "blonde thunder." Loki's
fly,
D. illusiolautus, means "elegant deception." Black
Widow's fly,
D. feminategus, means "woman wearing leather."
Deadpool's fly,
Humorolethalis sergius, is from Latin for
"wet and dead." Deadpool's fly shares his mask markings. (The
authors erected a new genus for Deadpool's fly, which previously was
known as
Daptolestes sergius (Walker).) See also
Stan Lee's
fly,
D. leei.
Eubetia boop Brown (tortricid moth)
Gekko hulk Grismer et al. 2022 (gecko)
referring to The Incredible Hulk, created by Stan Lee and Jack Kirby
in 1962. Like its namesake, the gecko becomes a large, green,
strong, aggressive beast when angry.
Hypocaccus kidpaddlei Gomy 2007 (histerid
beetle) Named for Kid Paddle, a Franco-Belgian comic, because the
beetle looks like a "blork", a moster from the Kid Paddle videogame
universe. [Nouvelle Revue d'Entomologie (N.S.), 24(2):
125]
Kalelia Pé & del Río,
2017 (Paleocene bivalve) "The name honors Kal-El, Kryptonian
name of Superman, the first superhero in comic books. This genus is
one of the oldest and largest alticostate carditids."
[J. Paleo. 69]
Kariridraco dianae Cerqueira et al. 2021
(Cretaceous pterosaur) Referring to Diana Prince, alter ego of Wonder
Woman.
Lepidopa luciae Boyko, 2002 (sand crab
(Crustacea: Anomura: Albuneidae)) Boyko originally thought to name it
for cartoonist Charles Schulz ("Peanuts") but Schulz's wife had the
bright suggestion to name it for Lucy van Pelt as her character was
known for being crabby. The monograph in which it is described
includes an appropriate "Peanuts" strip.
Nemesia shenlongi Pertegal et al. 2022
(trapdoor spider) Named after the wish-granting dragon Shenlong from
the Japanese Dragon Ball franchise.
Ninjemys Gaffney, 1992 (fossil turtle) Etymology:
"Ninja, in allusion to that totally rad, fearsome foursome epitomizing
shelled success; emys, turtle."
Scelio dupondi and
Scelio janseni Yoder, 2014 (parasitic
wasps) Named for the Thomson and Thompson twin detectives in Tintin
comics (Dupond and Dupont in French, Jansen and Janssen in Dutch).
The two were always being confused with one another.
Thanos Delcourt & Vidoi Iori, 2018
(theropod dinosaur) The genus is derived both from the Greek
word thanato, "death", and the Marvel Comics villain Thanos.
[Historical Biology, DOI:10.1080/08912963.2018.1546700]
Trigonopterus asterix, T. idefix, and
T. obelix Riedel, 2019 (weevils)
Named after characters from the French Asterix comics.
[ZooKeys 828:1]
Abaddon despoliator Derkarabetian, 2021
(opilionid) Refers to Abaddon the Despoiler, a characters from
Warhammer 40,000.
Aerodactylus Vidovic & Martill, 2014
(Jurassic pterosaur) Named after the Pokémon
Aerodactyl.
Binburrum articuno, B. moltres, and
B. zapdos Hsiao & Pollock, 2020
(beetles) Named after rare Pokemon, befitting the rarity of the
beetles.
Chilicola charizard Monckton, 2016 (bee)
Named after the Pokémon Charizard.
Cortana Salvador & Simone, 2013
(Paleocene land gastropod) Named after the character Cortana from the
HALO video-game series, alluding to markings on the shell surface.
[Papéis Avulsos de Zoologia 53: 16.]
Crash bandicoot Travouillon et al. 2014
(Miocene bandicoot relative) Named after the video game character
Crash Bandicoot, with an allusion to "a new radiation of more modern
bandicoots that 'crashed' through to dominate younger, drier
ecosystems of Australia." [J. Vert. Paleo. 34:
377.]
Demyrsus digmon Hsiao & Oberprieler,
2020 (weevil) Named after the Digimon Digmon; both the weevil
and the digital monster have an ability to drill.
Euconnus hosakae Hoshina, Fukutomi, &
Watanabe, 2020 (rove beetle) Dedicated to Miss Miyuki Hosaka,
heroine of the Japanese computer game Sentimental Graffiti;
she is from the type locality, Kanazawa City.
Galagadon Gates et al. 2019 (Cretaceous
shark) Named after the 1980s video game Galaga because the shark's
stepped triangular teeth resembled spaceships in that
game.
Halystina umberlee Salvador, Cavallari &
Simone, 2014 (gastropod) Named after a deep sea goddess from the
Faerûnian pantheon in the Forgotten Realms campaign setting of
the Dungeons and Dragons role-playing game. [Zootaxa 3878:
536.]
Rapturella ryani Salvador & Cunha, 2016
(gastropod) Named for Rapture, the deep-sea city from the
science-fiction video game BioShock, and Andrew Ryan, its founder.
[J. Molluscan Studies 6-9.]
Rotundicardia mariobrosorum Pé &
del Río, 2017 (Paleocene bivalve) "The specific epithet
honors Mario and Luigi, the Mario Bros. brothers, main characters
from the popular videogame Mario Bros., in which they collect
mushrooms, and it is a reference to the 'funginate' nodes of the
radial ribs in this species." [J. Paleo. 69]
Stentorceps weedlei Nielsen &
Buffington (wasp) Named for the Weedle, a hymenopteran larva
from the Pokémon video game series. "Weedle shares the
distinguishing character of S. weedlei, a spine in the middle of its
head."
Bidenichthys beeblebroxi Paulin, 1995 (triple-fin blenny)
with a false head pattern.
Erechthias beeblebroxi Robinson & Nelson, 1993 (tineid)
with a false head; after Zaphod Beeblebrox, two-headed character from
Douglas Adams' Hitchhiker's Guide to the Galaxy.
Babelichthys Davesne, 2017 (Eocene
crestfish) Named for the Babelfish from Douglas Adams' books mostly
because it is weird-looking.
Fiordichthys slartibartfasti Paulin, 1995 (brotulid, a
deep-sea fish) Named for Hitchhiker's Guide character
Slartibartfast, who is noted for designing fjords.
Medusaceratops lokii Ryan, Russell & Hartman,
2010 (Cretaceous ceratopsan) Named after the Medusa
specifically from Ray Harryhausen's "Clash of the Titans", for the
"snake-like, rocky appendages coming out the back of its skull"; and
after the Loki supervillian from Marvel comics, who was drawn with a
helmet with two giant curved horns.
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